INTERNATIONAL HERBICIDE-RESISTANT WEED DATABASE
select
Weeds Resistant to PS I Electron Diversion HRAC Group 22 (Legacy D)
by species and country
Weeds Resistant to PS I Electron Diversion HRAC Group 22 (Legacy D)
by species and country
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Weeds Resistant to PS I Electron Diversion HRAC Group 22 (Legacy D)
by species and country
#SpeciesCountryFirst Year
1 Alopecurus japonicus
Japanese Foxtail 		2010 - China
20105Alopecurus japonicusJapanese FoxtailD722PS I Electron Diversion
2 Amaranthus blitum (ssp. oleraceus)
Livid Amaranth 		1990 - Malaysia
199013Amaranthus blitum (ssp. oleraceus)Livid AmaranthD722PS I Electron Diversion
3 Arctotheca calendula
Capeweed 		1986 - Australia (Victoria)
198621Arctotheca calendulaCapeweedD722PS I Electron Diversion
4 Bidens pilosa
Hairy Beggarticks 		1991 - Kenya
199132Bidens pilosaHairy BeggarticksD722PS I Electron Diversion
5 Convolvulus arvensis
Field Bindweed 		2011 - Jordan
2011312Convolvulus arvensisField BindweedD722PS I Electron Diversion
6 Conyza bonariensis
Hairy Fleabane 		1989 - Egypt
1989 - Japan
2003 - South Africa
2009 - United States (California) *Multiple - 2 SOA's
2016 - Australia (New South Wales )
2022 - Jordan *Multiple - 2 SOA's
 		198960Conyza bonariensisHairy FleabaneD722PS I Electron Diversion
7 Conyza canadensis
Horseweed 		1980 - Japan
1993 - Canada (Ontario)
1994 - United States (Mississippi)
1998 - Belgium
2003 - United States (Delaware)
2007 - United States (Mississippi) *Multiple - 2 SOA's
2014 - United States (California) *Multiple - 2 SOA's
2024 - United States (New York)
198061Conyza canadensisHorseweedD722PS I Electron Diversion
8 Conyza sumatrensis
Sumatran Fleabane 		1980 - Taiwan
1986 - Japan
1990 - Malaysia
1998 - Sri Lanka
2016 - Brazil
2017 - Brazil *Multiple - 3 SOA's
2017 - Brazil *Multiple - 5 SOA's
2017 - Paraguay *Multiple - 3 SOA's
2018 - Australia (Queensland)
2024 - Bolivia
198065Conyza sumatrensisSumatran FleabaneD722PS I Electron Diversion
9 Crassocephalum crepidioides
Redflower Ragleaf 		1990 - Malaysia
199066Crassocephalum crepidioidesRedflower RagleafD722PS I Electron Diversion
10 Cuphea carthagenensis
Tarweed Cuphea 		1984 - Fiji
198468Cuphea carthagenensisTarweed CupheaD722PS I Electron Diversion
11 Eleusine indica
Goosegrass 		1990 - Malaysia
1996 - United States (Florida)
2009 - Malaysia *Multiple - 2 SOA's
2009 - Malaysia *Multiple - 4 SOA's
2010 - China
2012 - Indonesia *Multiple - 2 SOA's
2012 - Indonesia
2015 - Australia (Queensland)
2016 - Colombia *Multiple - 2 SOA's
2019 - United States (Alabama)
199081Eleusine indicaGoosegrassD722PS I Electron Diversion
12 Epilobium ciliatum
Fringed Willowherb 		1982 - Belgium
1989 - United Kingdom
198282Epilobium ciliatumFringed WillowherbD722PS I Electron Diversion
13 Erigeron philadelphicus
Philadelphia Fleabane 		1980 - Japan
198085Erigeron philadelphicusPhiladelphia FleabaneD722PS I Electron Diversion
14 Gamochaeta pensylvanica
Pennsylvania Everlasting 		1991 - Japan
2015 - Australia (Queensland)
1991298Gamochaeta pensylvanicaPennsylvania EverlastingD722PS I Electron Diversion
15 Hedyotis verticillata
Woody borreria 		2005 - Malaysia *Multiple - 2 SOA's
 		2005310Hedyotis verticillataWoody borreriaD722PS I Electron Diversion
16 Hordeum murinum ssp. glaucum
Smooth Barley 		1984 - Australia (Victoria)
1990 - Australia (South Australia)
198497Hordeum murinum ssp. glaucumSmooth BarleyD722PS I Electron Diversion
17 Hordeum murinum ssp. leporinum
Hare Barley 		1988 - Australia (Victoria)
2001 - Australia (South Australia)
198898Hordeum murinum ssp. leporinumHare BarleyD722PS I Electron Diversion
18 Ischaemum rugosum
Saramollagrass 		1989 - Malaysia
198999Ischaemum rugosumSaramollagrassD722PS I Electron Diversion
19 Landoltia punctata
Dotted Duckweed 		2001 - United States (Florida)
2001232Landoltia punctataDotted DuckweedD722PS I Electron Diversion
20 Lepidium virginicum
Virginia Pepperweed 		1993 - Canada (Ontario)
1993103Lepidium virginicumVirginia PepperweedD722PS I Electron Diversion
21 Lolium perenne ssp. multiflorum
Italian Ryegrass 		2015 - United States (California) *Multiple - 3 SOA's
2016 - United States (California) *Multiple - 4 SOA's
2022 - United States (North Carolina) *Multiple - 4 SOA's
 		2015110Lolium perenne ssp. multiflorumItalian RyegrassD722PS I Electron Diversion
22 Lolium rigidum
Rigid Ryegrass 		2002 - South Africa
2003 - South Africa *Multiple - 3 SOA's
2010 - Australia (South Australia) *Multiple - 2 SOA's
2010 - Australia (South Australia) *Multiple - 5 SOA's
2013 - Australia (Western Australia) *Multiple - 2 SOA's
 		2002112Lolium rigidumRigid RyegrassD722PS I Electron Diversion
23 Mazus fauriei
Asian Mazus 		2011 - China
2011281Mazus faurieiAsian Mazus D722PS I Electron Diversion
24 Mazus pumilus
Japanese Mazus 		2006 - Japan
2006296Mazus pumilusJapanese MazusD722PS I Electron Diversion
25 Mitracarpus hirtus
Tropical Girdlepod 		2007 - Australia (Queensland)
2007240Mitracarpus hirtus Tropical GirdlepodD722PS I Electron Diversion
26 Plantago lanceolata
Buckhorn Plantain 		2022 - South Africa *Multiple - 2 SOA's
 		2022220Plantago lanceolataBuckhorn PlantainD722PS I Electron Diversion
27 Poa annua
Annual Bluegrass 		1981 - United Kingdom
1993 - Belgium
1981130Poa annuaAnnual BluegrassD722PS I Electron Diversion
28 Sclerochloa dura
Hardgrass 		2011 - China
2011283Sclerochloa duraHardgrassD722PS I Electron Diversion
29 Solanum americanum
American Black Nightshade 		1985 - United States (Florida)
2009 - New Zealand
1985161Solanum americanumAmerican Black NightshadeD722PS I Electron Diversion
30 Solanum nigrum
Black Nightshade 		1990 - Malaysia
2009 - New Zealand
2015 - Australia (Queensland)
1990162Solanum nigrumBlack NightshadeD722PS I Electron Diversion
31 Solanum ptycanthum
Eastern Black Nightshade 		2009 - Canada (Ontario)
2009190Solanum ptycanthumEastern Black NightshadeD722PS I Electron Diversion
32 Vulpia bromoides
Squirreltail Fescue 		1990 - Australia (Victoria)
1990173Vulpia bromoidesSquirreltail FescueD722PS I Electron Diversion
33 Youngia japonica
Asiatic Hawksbeard 		1986 - Japan
2009 - China
1986175Youngia japonicaAsiatic HawksbeardD722PS I Electron Diversion
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Heap, I.  The International Herbicide-Resistant Weed Database.  Online.  Friday, October 31, 2025 .  Available  www.weedscience.org Copyright © 1993- 2025 WeedScience.org All rights reserved. Fair use of this material is encouraged. Proper citation is requested.